All

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 E05 Investigative Techniques D008919 +
  E05.196 Chemistry, Analytical D002623 +
   E05.196.181 Chromatography D002845 +
    E05.196.181.400 Chromatography, Liquid D002853 +
     E05.196.181.400.454 Chromatography, Paper D002854 +
      E05.196.181.400.454.655 Nucleotide Mapping D009710 +
        E05.196.181.400.454.655.150 Blotting, Southern D015139 
        E05.196.181.400.454.655.100 Blotting, Northern D015152 
 E05 Investigative Techniques D008919 +
  E05.196 Chemistry, Analytical D002623 +
   E05.196.401 Electrophoresis D004586 +
    E05.196.401.319 Electrophoresis, Paper D004590 +
     E05.196.401.319.670 Nucleotide Mapping D009710 +
       E05.196.401.319.670.150 Blotting, Southern D015139 
       E05.196.401.319.670.100 Blotting, Northern D015152 
 E05 Investigative Techniques D008919 +
  E05.196 Chemistry, Analytical D002623 +
   E05.196.680 Nucleotide Mapping D009710 +
     E05.196.680.150 Blotting, Southern D015139 
     E05.196.680.100 Blotting, Northern D015152